分类： 生物学 >> 生物进化论 提交时间： 2024-03-15
摘要： The previously alleged ‘eugnathid amiiform’ Sinoeugnathus kueichowensis is asmall-sized halecomorph from the Middle Triassic (Ladinian) marine deposits of Guizhou andYunnan, China. A morphological redescription and taxonomic revision of this taxon are providedbased on a detailed examination of 15 new specimens. Among them, IVPP V24315 (standardlength = 64 mm) is appointed as the neotype, given that the holotype is missing. Studies of thesespecimens revealed some morphological details previously undescribed or misidentified forthis taxon, including a hatchet-shaped antorbital, two broad suborbitals, a sensory canal in themaxilla, and three pairs of extrascapulars. For the first time, Sinoeugnathus was incorporatedinto an analysis of halecomorph phylogeny, and the results recover it as the sister taxon of theAnisian Subortichthys from Luoping, Yunnan, and both are grouped with two Ladinian generaAllolepidotus and Eoeugnathus from the Monte San Giorgio area into a monophyletic group(namely Subortichthyidae fam. nov. herein) at the base of Ionoscopiformes. This taxonomicreassessment of Subortichthys provides new insights into the phylogeny and paleogeographicevolution of Ionoscopiformes.
分类： 生物学 >> 动物学 提交时间： 2017-11-07 合作期刊: 《古脊椎动物学报》
摘要： Ionoscopiformes, sister to Amiiformes, are an extinct group of halecomorph fishes previously known mainly from the late Middle Triassic (Ladinian) and Late Jurassic of Europe and the Early Cretaceous of the New World. They have not been known in China until recently two ionoscopiforms (Robustichthys and Panxianichthys) were found from the early Middle Triassic (Anisian) of Yunnan and Guizhou, respectively. Here, a new ionoscopiform, Subortichthys triassicus gen. et sp. nov., is described on the basis of six well-preserved specimens from the early Middle Triassic (Pelsonian, Anisian, ~244 Ma) marine deposits exposed in Luoping, eastern Yunnan, China. The discovery documents the second ionoscopiform from the Luoping fossil beds and one of the oldest ionoscopiforms in the world, providing important information for understanding the origin and early diversification of this group. Subortichthys is an unambiguous halecomorph as it possesses two synapomorphies of this clade: a symplectic articulating with the lower jaw and a notched posterior margin of the maxilla. Within the phylogenetic framework of the Halecomorphi, results of the phylogenetic analysis place Subortichthys at the base of the Ionoscopiformes because it bears a sensory canal in the maxilla (an ionoscopiform synapomorphy), but lacks derived features of other ionoscopiforms. Notably, Subortichthys is distinguished from other members of this order with a unique combination of features, such as presence of three or four pairs of extrascapulars and a much expanded third infraorbital posteriorly contacting the preopercle. The new material reveals that the earliest diversification of ionoscopiforms was well undertaken in South China (a part of east Paleotethys Ocean then) by the early Middle Triassic (Anisian).
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